Evan MacLean

About Evan MacLean

I am an evolutionary anthropologist and comparative psychologist with research interests in how and why cognition evolves.  My research program is motivated both by questions about what makes the human mind unique, and broader evolutionary questions regarding the proximate mechanisms and functional significance of cognition.  Studies in my lab address cognition within an explicitly phylogenetic comparative framework, and integrate ideas and methods from cognitive science and evolutionary biology.  

Selected Publications

For a complete list of publications from our lab please see our google scholar page.

MacLean, E. L., Gesquiere, L., Gruen, M., Sherman, B., Martin, W. L., & Carter, C. S. (2017). Endogenous Oxytocin, Vasopressin and Aggression in Domestic Dogs. Frontiers in Comparative Psychology.

MacLean, E. L., Gesquiere, L., Gee, N., Levy, K., Martin, W. L., & Carter, C. S. (2017). Effects of Human-Animal Interaction on Dog Salivary and Plasma Oxytocin and Vasopressin. Frontiers in Comparative Psychology.

MacLean, E. L. & Nunn, C. L. (2017) Phylogenetic Approaches for Research in Comparative Cognition.  In J. Call, G. Burghardt, I. Pepperberg, C. Snowdon & T. Zentall (Eds.), APA Handbook of Comparative Psychology. American Psychological Association.

MacNamara, M.  & MacLean, E. L. Selecting Animals for Education Environments.  (2017). In N. Gee, A. Fine, & P. McCardle (Eds.) How Animals Help Students Learn: Research and Practice for Educators and Mental Health Professionals. Routledge, Taylor Francis Group.  

Lucca, K., MacLean, E. L., & Hare, B. (2017). The development and flexibility of gaze alternations in bonobos and chimpanzees. Developmental Science

MacLean, E. L., Gesquiere, L. R., Gee, N., Levy, K., Martin, W. L., & Carter, C. S. (2017). Validation of Salivary Oxytocin and Vasopressin as Biomarkers in Domestic Dogs. Journal of Neuroscience Methods

MacLean, E. L., Herrmann, E., Suchindran, S., & Hare, B. (2017). Individual differences in cooperative communicative skills are more similar between dogs and humans than chimpanzees. Animal Behaviour, 126, 41-51.

Brandtzaeg, O. K., Johnsen, E., Roberg-Larsen, H., Seip, K. F., MacLean, E. L., Gesquiere, L. R., Leknes, S., Lundanes, E. & Wilson, S. R. (2016). Proteomics tools reveal startlingly high amounts of oxytocin in plasma and serum. Scientific Reports, 6.

Reddy, R. B., Krupenye, C., MacLean, E. L., Hare, B. (2016). No evidence for contagious yawning in lemurs.  Animal Cognition.

MacLean, E. L. (2016). Unravelling the Evolution of Uniquely Human Cognition.  Proceedings of the National Academy of Sciences.

Krupenye, C., MacLean, E. L., & Hare, B. (2016). Does the bonobo have a (chimpanzee-like) theory of mind? Bonobos: Unique in mind, brain and behavior: Oxford University Press.

MacLean, E. L. & Hare, B. (2016). Comparative Perspectives on Lemur Cognition. In E. Palagi & I. Norscia (Eds.), The Missing Lemur Link: An Ancestral Step in Human Evolution. Cambridge University Press.

Stewart, L., MacLean, E. L., Ivy, D., Woods, V., Cohen, E., Rodriguez, K., McIntyre, M., Mukherjee, S., Call, J., Kaminski, J., Miklósi, A.,  Wrangham, W. & Hare, B. (2015). Citizen science as a new tool in dog cognition research. PloS one, 10(9), e0135176.

MacLean, E. L., & Hare, B. (2015). Dogs hijack the human bonding pathway. Science, 348(6232), 280-281.

Bray, E. E., MacLean, E. L., & Hare, B. (2015). Increasing arousal enhances inhibitory control in calm but not excitable dogs. Animal Cognition.

Reddy, R., MacLean, E. L., Sandel, A. A., & Hare, B. (2015). Social and nonsocial inhibitory control in five lemur species. Primates, (56), 241-252.

MacLean, E. L. & Hare, B. (2015). Bonobos and chimpanzees exploit helpful but not prohibitive gestures. Behaviour, 152(3-4), 493-520.

MacLean, E. L., Hare, B., Nunn, C. L., Addessi, E., Amici, F., Anderson, R. C., Aureli, F., Baker, J. M., Bania, A. E., Barnard, A. M., Boogert, N. J., Brannon, E. M., Bray, E. E., Bray J., Brent, L. J. N., Burkart, J. M., Call J., Cantlon, J. F., Cheke, L. G., Clayton, N. S., Delgado, M. M., DiVincenti, L. J., Fujita, K., Herrmann, E., Hiramatsu, C., Jacobs, L. F., Jordan, K. E., Laude, J. R., Leimgruber, K. L., Messer, E. J. E. , Moura, A. C. de A., Ostojic, L., Picard, A., Platt, M. L., Plotnik, J. M., Range, F., Reader, S. M., Reddy, R. B., Sandel, A. A., Santos, L. R., Schumann, K., Seed, A. M., Sewall, K. B., Shaw, R. C., Slocombe, K. E., Su, Y., Takimoto, A., Tan, J., Tao, R., van Schaik, C. P., Virányi, Z., Visalberghi, E., Wade, J. C., Watanabe, A., Widness, J., Young, J. K., Zentall, T. R., & Zhao, Y. (2014). The evolution of self-control. Proceedings of the National Academy of Sciences, 111 (20), E2140-E2148.

MacLean, E. L., Krupenye, C., & Hare, B. (2014). Dogs account for body orientation but not visual barriers when responding to pointing gestures. Journal of Comparative Psychology, 128(3), 285-197.

Bray, E., MacLean, E. L., & Hare, B. (2014). Context-specificity of inhibitory control in dogs. Animal Cognition. 17(1), 15-31.

MacLean, E. L., Sandel, A. A., Bray, J., Oldenkamp, R., Reddy, R., & Hare, B. (2013). Group size predicts social but not nonsocial cognition in lemurs. PLOS ONE, 8(6),e66359.

MacLean, E. L. & Hare, B. (2013). Spontaneous triadic engagement in chimpanzees and bonobos. Journal of Comparative Psychology. 127(3), 245-255

MacLean, E. L., & Hare, B. (2012). Bonobos and chimpanzees infer the target of another's attention. Animal Behaviour, 83(2), 345-353. 

MacLean E. L., Matthews L., Hare B., Nunn C., Anderson R., Aureli F., Brannon E., Call J., Drea C., Emery N., Haun D., Herrmann E., Jacobs L., Platt M., Rosati A., Sandel A., Schroepfer K., Seed A., Tan J., van Schaik C., & Wobber V. (2012). How does cognition evolve? Phylogenetic comparative psychology. Animal Cognition, 15(2), 1-16.

MacLean, E. L., Mandalaywala, T., & Brannon, E. (2012). Variance-sensitive choice in lemurs: Constancy trumps quantity. Animal Cognition, 15(1), 1-11.

Sandel, A. A., MacLean, E. L., & Hare, B. (2011). Evidence from four lemur species that ringtailed lemur social cognition converges with that of haplorhine primates. Animal Behaviour, 81(5), 925-931.

Merritt, D., MacLean, E. L., Crawford, J. C., & Brannon, E. M. (2011). Numerical rule-learning in ring-tailed lemurs (Lemur catta). Frontiers in Comparative Psychology, 2.

MacLean, E. L., Barrickman, N. L., Johnson, E. M., & Wall, C. (2009). Sociality, ecology, and relative brain size in lemurs. Journal of Human Evolution, 56(5), 471-478.

MacLean, E. L., Prior, S. R., Platt, M. L., & Brannon, E. M. (2009).  Primate location preference in a double-tier cage: The effects of illumination and cage height. Journal of Applied Animal Welfare Science, 12(1), 73-81.

MacLean, E. L., Merritt, D. J., & Brannon, E. M. (2008). Social complexity predicts transitive reasoning in prosimian primates. Animal Behaviour, 76(2), 479-486

Jordan, K. E., MacLean, E. L., & Brannon, E. M. (2008). Monkeys match and tally quantities across senses. Cognition, 108(3), 617-625.

Merritt, D. J., MacLean, E. L., Jaffe, S., & Brannon, E. M. (2007).  A comparative analysis of serial ordering in ring-tailed lemurs (Lemur catta). Journal of Comparative Psychology, 121(4), 363-371.



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Contact Information

Assistant Professor
Telephone: 520.621.0386
Office: 313A2


Ph.D. (Evolutionary Anthropology) – Duke University

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