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Daniel Horschler

About Daniel Horschler

I am a Ph.D. candidate in Biological Anthropology working with Dr. Evan MacLean. I graduated from the University of North Carolina at Chapel Hill in 2016 with a B.S. in Psychology and minor in Anthropology, and I received an M.A. in Anthropology from the University of Arizona in 2018.

Prior to joining the School of Anthropology, I worked in a number of diverse research roles, including projects examining human pain perception, metacognition, and memory at UNC, user experience (UX) at Lenovo, and canine and non-human primate cognition in Yale University’s Canine Cognition Center and with Yale’s Comparative Cognition Laboratory at the Cayo Santiago Biological Field Station in Puerto Rico.

My research interests broadly center on cognitive evolution, with a particular focus on sociocognitive abilities relating to theory of mind and shared intentionality. I am also interested in relationships between neuroanatomy and cognition both within and across species. I investigate these topics by integrating non-invasive cognitive, behavioral, and biological research in dogs, wolves, and free-ranging non-human primates.

My research has been covered by national and international media outlets including SciencePsychology Today, ScienceDaily, InsiderSky News, and talkRADIO London as well as by local media outlets including Arizona Public MediaUA NewsThe Daily WildcatCBS13 Tucson, ABC15 PhoenixKVOI Tucson RadioKJZZ Phoenix Radio, and StarTribune. I have also commented on related research in popular science articles for National Geographic, Science, and The Washington Post.

I am currently affiliated with the Cognitive Science Graduate Interdisciplinary Program, and have worked as a Graduate Teaching Associate in both the School of Anthropology and the School of Mind, Brain and Behavior.

Click here to view my full CV

Selected Publications

Salomons, H., Smith, K., Callahan-Beckel, M., Callahan, M., Levy, K., Kennedy, B.S., Bray, E., Gnanadesikan, G.E., Horschler, D.J., Gruen, M., Tan, J., White, P., MacLean, E., & Hare, B. (Preprint). Cooperative communication with humans evolved to emerge early in dogs. bioRxiv

Bray, E.E., Gnanadesikan, G.E., Horschler, D.J., Levy, K.M., Kennedy, B.S., Famula, T.R., & MacLean, E.L. (In press). Early-emerging and highly-heritable sensitivity to human communication in dogs. Current Biology.

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2021). How do non-human primates represent others’ awareness of where objects are hidden? Cognition.

Arre, A.M.* & Horschler, D.J.* (2021). Swimming and diving as social play in juvenile rhesus macaques (Macaca mulatta). Behaviour. [*Denotes equal contribution].

Bray, E.E., Gruen, M.E., Gnanadesikan, G.E., Horschler, D.J., Levy, K.M., Kennedy, B.S., Hare, B.A., & MacLean, E.L. (2020). Dog cognitive development: A longitudinal study across the first two years of life. Animal Cognition, 24(2), 311-328.

Horschler, D.J., MacLean, E.L., & Santos, L.R. (2020). Advancing gaze-based research on primate theory of mind. Trends in Cognitive Sciences, 24(10), 778-779.

Bray, E.E., Gruen, M.E., Gnanadesikan, G.E., Horschler, D.J., Levy, K.M., Kennedy, B.S., Hare, B.A., & MacLean, E.L. (2020). Cognitive characteristics of 8-to-10-week-old assistance dog puppies. Animal Behaviour, 166, 193-206.

Horschler, D.J., MacLean, E.L., & Santos, L.R. (2020). Do non-human primates really represent others' beliefs? Trends in Cognitive Sciences, 24(8), 594-605.

Horschler, D.J. & MacLean, E.L. (2019). Leveraging brain-body scaling relationships for comparative studies. Animal Cognition, 22(6), 1197-1202.

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2019). Do non-human primates really represent others' ignorance? A test of the awareness relations hypothesis. Cognition, 190, 72-80.

Horschler, D.J., Hare, B., Call, J., Kaminski, J., Miklósi, Á., & MacLean, E.L. (2019). Absolute brain size predicts dog breed differences in executive function. Animal Cognition, 22(2), 187-198.

Courses Taught

Graduate Teaching Associate: Fundamentals of Neuroscience and Cognitive Science | Dog Thought | Animal Minds

Graduate Teaching Assistant: Research Methods in Cognitive Science | Human Variation in the Modern World

Areas of Study

Cognitive Evolution | Cognitive Development | Social Cognition | Theory of Mind | Joint Experience | Shared Intentionality | Executive Function | Domestication | Brain Evolution | Neuroanatomy

Daniel Horschler's picture

Contact Information

Ph.D. Candidate, Haury Fellow
Office: Haury 313A3


M.A. - Anthropology - University of Arizona (2018)

B.S. - Psychology - University of North Carolina at Chapel Hill (2016)

Dissertation Title

The evolutionary origins of representational and motivational hallmarks of human social cognition